101 - Document - Document and Entity Information
link:calculationLink
link:presentationLink
link:definitionLink
103 - Statement - Condensed Consolidated Balance Sheets
link:calculationLink
link:presentationLink
link:definitionLink
104 - Statement - Condensed Consolidated Balance Sheets (Parenthetical)
link:calculationLink
link:presentationLink
link:definitionLink
105 - Statement - Condensed Consolidated Statements of Operations
link:calculationLink
link:presentationLink
link:definitionLink
106 - Statement - Condensed Consolidated Statements of Cash Flows
link:calculationLink
link:presentationLink
link:definitionLink
107 - Disclosure - Organization
link:calculationLink
link:presentationLink
link:definitionLink
108 - Disclosure - Summary of Significant Accounting Policies
link:calculationLink
link:presentationLink
link:definitionLink
109 - Disclosure - Property and Equipment
link:calculationLink
link:presentationLink
link:definitionLink
110 - Disclosure - Share-Based Compensation
link:calculationLink
link:presentationLink
link:definitionLink
111 - Disclosure - Collaborations and License Agreements
link:calculationLink
link:presentationLink
link:definitionLink
112 - Disclosure - Summary of Significant Accounting Policies (Policies)
link:calculationLink
link:presentationLink
link:definitionLink
113 - Disclosure - Summary of Significant Accounting Policies (Tables)
link:calculationLink
link:presentationLink
link:definitionLink
114 - Disclosure - Property and Equipment (Tables)
link:calculationLink
link:presentationLink
link:definitionLink
115 - Disclosure - Share-Based Compensation (Tables)
link:calculationLink
link:presentationLink
link:definitionLink
116 - Disclosure - Organization - Additional Information (Detail)
link:calculationLink
link:presentationLink
link:definitionLink
117 - Disclosure - Summary of Significant Accounting Policies - Additional Information (Detail)
link:calculationLink
link:presentationLink
link:definitionLink
118 - Disclosure - Summary of Significant Accounting Policies - Common Stock Equivalents Excluded in Calculation of Diluted Loss Per Share (Detail)
link:calculationLink
link:presentationLink
link:definitionLink
119 - Disclosure - Property and Equipment - Property and Equipment (Detail)
link:calculationLink
link:presentationLink
link:definitionLink
120 - Disclosure - Property and Equipment - Additional Information (Detail)
link:calculationLink
link:presentationLink
link:definitionLink
121 - Disclosure - Share-Based Compensation - Additional Information (Detail)
link:calculationLink
link:presentationLink
link:definitionLink
122 - Disclosure - Share-Based Compensation - Total Stock-Based Compensation Expense (Detail)
link:calculationLink
link:presentationLink
link:definitionLink
123 - Disclosure - Share-Based Compensation - Weighted-Average Assumptions Used for Calculating Value of Options Granted (Detail)
link:calculationLink
link:presentationLink
link:definitionLink
124 - Disclosure - Share-Based Compensation - Stock Options Activity (Detail)
link:calculationLink
link:presentationLink
link:definitionLink
125 - Disclosure - Collaborations and License Agreements - Additional Information (Detail)
link:calculationLink
link:presentationLink
link:definitionLink